PLANT COMMUNITIES OF BROAD-LEAVED FORESTS OF THE COLCHIS LOWLAND (WESTERN TRANSCAUCASIA)
https://doi.org/10.36305/0513-1634-2021-138-39-49
Abstract
Three associations of the class Carpino-Fagetea Jakucs et Passarge 1968 were described as а result of the classification of forest communities of the Colchis Lowland (Westem Transcaucasia) using the Brown Blanquet method. Association Vaccinio arctostaphilli-Carpinetum betuli ass. nova prov. was included in the order Lathyro-Carpinetalia Passarge 1981, alliance Crataego-Carpinion Passarge 1981. Associations Truello thunbergii-Alnetum barbatae ass. nova prov. and Carici remotae-Pterocmyetum pterocarpae ass. nova prov. were included in the order Rhododendro ponticae-Fagetalia orientalis Quezel et al. 1992, alliance Alnion barbatae Quezel et al. 1992. The results of the DCA ordination and the interpretation of the leading axes demonstrated three well-distinguished ecological and floristic types of deciduous forests (ofthe association rank) forming the ecological series according to the leading ecological factors - the degree of humidity of habitats and the degree of drainage of substrates. Twenty-three plant species - thermophilous Tertiary rehcs were found in all three associations. Of these, eight evergreen species play а significant phytocoenotic role in the formation of the understory in forest communities.The pecuharities of the Tertiary rehc speciesdistributions in the floristic compositions of described associations are discussed.
Keywords
About the Authors
N. B. ErmakovRussian Federation
Yu. V. Plugatar
Russian Federation
V. D. LeiЬa
Russian Federation
E. V. Ermakova
Russian Federation
References
1. Буш НА. Ботаника-географический очерк Европейской части СССР и Кавказа. -М.-Л.: Изд-во АН СССР, 1936. - 327 с.
2. Гроссгейм А.А. Растительный покров Кавказа. - М.: Изд-во МОИП, 1948. - 153с.
3. Малеев В.П. Третичные реликты во флоре Западного Кавказа и основные этапы четвертичной истории его флоры и растительности // Материалы по истории флоры и растительности СССР. -М.-Л.: Изд.-во АН СССР, 1941. -Вып. 1. -С. 61-144.
4. Гулисашвили В.З., Махатадзе ЛБ., Прилипко ЛИ. Растительность Кавказа. - М.: Наука, 1975. -233 с.
5. Поварницын В.А. Типы лесов Абхазии // Под ред. Келлера Б.А. Абхазия. Геоботанический и лесоводческий очерк. По материалам экспедиции Академии наук СССР в 1934 г. -М.-Л.: Изд-во АН СССР, 1936. - С. 67-156.
6. Гребенщиков О.С., Шанина А.А., Белоновская Е.А. 1990. Леса крайней западной части Большого Кавказа// Биота экосистем Большого Кавказа.- М.- С. 63-84.
7. Passarge Н Carpineta im kartalinischen Kaukasus // Phytocoenologia. - 1981. - Vol. 9(4 ). - Р. 533-545. DOI: https://doi.org/10.1002/fedr.19810920507
8. Novak Р., Zukal D., Kalnikova V, Chytry К., Kavgaci А. Ecology and syntaxonomy of Colchic forests in south-western Georgia (Caucasus region) // Phytocoenologia. - 2019. - Vol. 49 (3 ). - Р. 231-248. DOI: https://doi.org/10.l127/phyto/2019/0250
9. Hennekens S.M, Schaminee JHJ Turboveg, а Comprehensive Data Base Management System for Vegetation Data. Journal of Vegetation Science. - 2001. - Vol. 12. - Р. 589-591. DOI: https://doi.org/10.2307/3237010
10. Тichy L. JШСЕ. Software for Vegetation Classification // Journal of Vegetation Science. - 2002. - Vol. 13. - Р. 451-453. DOI: https://doi.org/10.llll/j.1654-l103.2002.tb02069.x
11. Westhoff V, Maarel Е. Тhе Braun-Blanquet Approach // In: Ordination and Classification of Communities (R.H Whittaker, ed.). - Dordrecht: Dr. W. Junk,1973. - Р. 617-626.
12. Миста L., Bйltmann Н, DierJJen К., Theurillat JP., Raus Т, Сат! А., Sumberova К., Willner W, Dengler J, Garcia R, Chytry М, Hajek М, Di Pietro R, Iakushenko D., Pallas J, Daniёls F., Bergmeier Е., Guerra А., Ermakov N., Valachovic М, Schaminee J, Lysenko Т, Didukh У, Pignatti S., Rodwell J, Саре/о J, Weber Н, Solomeshch А., Dimopoulos Р., Aguiar С., Hennekens S., Тichy L. Vegetation of Europe: hierarchical floristic classification systemof vascular plant, bryophyte, lichen, and algal communities // Applied Vegetation Science. - 2016. - Vol. 19 (1). - Р. 3-264. DOI: 10.1111/avsc.12257.
13. СоЬап S., Willner W Numerical classification of the forest vegetation in the Western Euxine Region of Turkey // Phytocoenologia -2019. - Vol. 49 (1). - Р. 71-106. DOI:10.l127/phyto/2018/0274
14. Bergmeier Е., Walentowski Н, Gйngдroglu С. Turkish Forest Habitat Types - An Annotated Conspectus Based оп the EU Habitats Directive with Suggestions for an Upgrade // In: Practicability of EU Natura 2000 Concept in the Forested Areas of Turkey.- Ankara: Turkey Foresters' AssociationPuЫ., 2019. - Р. 134-292.
15. Novak Р., Willner W, Zukal D., кьи» J, Rolecek J, Swierkosz К., Ewald J, Wohlgemuth Т, Csiky J, Onyshchenko V, Chytry М Oak-hornbeam forests of central Europe: а formalized classification and syntaxonomic revision // Preslia. - 2020. - Vol. 92. - Р. 1-34. DOI: 10.23855/preslia.2020.001
16. Theurillat JP., Willner W, Еетапаег-Сопгагег F., Bueltmann Н, Сапи А., Gigante D., Миста L., Weber НЕ. International Code of Phytosociological Nomenclature. 4th edition // Applied Vegetation Science. - 2020. - V. 24(2). - Р. 1-62. DOI: https://doi.org/10.1111/avsc.12491.
17. Czerepanov S.К. Vascular plants ofRussia and adjacent states (the former USSR). - Cambridge: Cambridge UniversityPress,1995. - 516 р.
Review
For citations:
Ermakov N.B., Plugatar Yu.V., LeiЬa V.D., Ermakova E.V. PLANT COMMUNITIES OF BROAD-LEAVED FORESTS OF THE COLCHIS LOWLAND (WESTERN TRANSCAUCASIA). Bulletin of the State Nikitsky Botanical Gardens. 2021;1(138):39-49. (In Russ.) https://doi.org/10.36305/0513-1634-2021-138-39-49